Guioa pterorhachis Welzen, Blumea 33: 419 (1988)

Latin for 'winged stalks', referring to the winged leaf rachis.


Shrub to tree, 3-16 m high, dbh 10 cm to 1.15 m. Branchlets glabrous, at most sparsely hairy when young. Leaves 3-7-jugate; rachis winged, wing up to 4 mm broad, (sub)glabrous, petiole 1.1-12.2 cm long. Leaflets subsessile, opposite to subopposite, elliptic (to obovate), 2.4-14.6 by 1-4.6 cm, index 1.9-3.9, often slightly asymmetrical, especially basally, (sub)coriaceous, punctate; base acute to attenuate to cuneate; margin entire, flat; apex gradually acuminate to cuspidate; upper surface glabrous (to puberulous) on the midrib and venation; lower surface dull, papillate, (very) sparsely, usually shortly sericeous, domatia many sunken sacs in axils of nerves; venation on upper surface (slightly sunken to) flat to raised, raised on lower; midrib below hardly raised, flat; nerves 0.2-3.6 cm apart, marginally looped and joined, less so in lower half of leaflets; nerves laxly reticulate, rather indistinct. Inflorescences axillary (to pseudoterminal), (unbranched to) branching basally and along the terete, subsericeous, 1-20.8 cm long axis; first order branches up to 11.5 cm long; cymules cincinnate, 2-5-flowered; bracts and bracteoles deltate to triangular, outside sericeous, inside (sub)glabrous; bracts 0.7-1.2 mm long; bracteoles 0.3-0.8 mm long; pedicels 2-7 mm long, sericeous. Flowers 3.5-4 mm in diam. Sepals 5. ovate, margin (and outside) pilose, margin with glands, inside (sub)glabrous; 2 outer smaller ones 1-1.9 by 0.8-1.3 mm: 3 inner larger ones 1.7-3 by 1.2-2.7 mm, margin petaloid. Petals 5, elliptic (to obovate), 1.1-2.8 by 0.8-1.6 mm, white; claw 0.2-0.5 mm high; margin pilose, outside and inside glabrous, apex acute; scales 1.1-2 mm long, free; crest a pilose flat part of bifid scale apex. Disc interrupted to uninterrupted. Stamens 8; filaments 2-5 mm long, pilose, especially basally; anthers c. 0.3 mm long, glabrous. Pistil: ovary 0.3-1 mm long, sparsely hirsute; style and stigma 0.1-1 mm long. Fruits with 1-3 well developed lobes, 1.4-1.9 by 1.2-2.2 cm. (smooth to) somewhat rugose-ruminate. glabrous, yellow to red when fresh, blackish when dry; stipe 4-7 mm high, slender; margin blunt; lobes 8.5-1 1 by 8-12 mm. Seeds obovoid, 7.5-10.5 by 6-9 mm; hilum 1.7-2 mm long; arillode edible, with yellow exudate. [From Flora Malesiana]

In understorey of primary and especially secondary forest, along rivers and roadsides, on flat to undulating country. Soil: white, yellow, or black sands; altitude sea level up to 30 (-500) m.



Local names