Aporosa lucida (Miq.) Airy Shaw, Kew Bull. Add. Ser. 4 (1975)

Synonyms
Aporosa borneensis Pax & K.Hoffm.
Aporosa microsphaera Hook.f.
Aporosa miqueliana Mull.Arg.
Tetractinostigma lucidum Miq.

Diagnostics
Understorey tree op to 18 m tall and 16(-25) cm dbh. Stipules ca. early dropped. Leaves alternate, simple, penni-veined, glabrous. Flowers ca. 0.3 mm in diameter, yellow, placed in racemes. Fruits ca. 8 mm long, yellow-orange-red-purplish, dehiscent capsules, seed with orange aril.

Description
(Small) tree or shrub, up to 18 m high, up to 25 cm diameter. Bark (pale or dark) brown, (pale) fawn, (pale) grey, fawn-grey, reddish yellow, yellowish, whitish, or greenish, grey mottled, dippled, smooth, scaly, or lenticellate to rugose, thin, papery flaky; inner bark (pale) brown, (pale) yellow, yellowish brown, red-brown, or greenish brown, thin, fibrous. Sapwood (pale) brown, (pale) yellow, yellowish, light orange, or white, hard. Young branches white, very sparsely puberulous. Stipules not seen, caducous. Petiole half-terete, ridged, 6-25 by 0.8-1.5 mm, glabrous, lower pulvinus 1.5-3 by 1-2 mm, upper pulvinus 2-5 by 1-2.2 mm, both distinct. Leaves narrowly ovate to narrowly obovate, 5.5-24 by 2-9 cm; base obtuse to attenuate, basal glands present, fading; margin lowly undulate to lowly glandular-crenate, marginal glands regular, distinct, brownish; apex acuminate to cuspidate; thin, smooth, sometimes brittle, rather shiny, drying yellow-brown to dark green above and beneath; completely glabrous; dots rather laxly set, fine, greyish to black, fading; disc-like glands small, few, scattered within marginal arches, fading. Nervation: midrib prominent on both sides; nerves 8-11 pairs, slightly prominent to flat above, prominent beneath, marginal arches distinct, 1.5-6 mm from the margin; tertiary veins and venation distinct, fading in older leaves, densely reticulate, slightly prominent to flat. Inflorescences axillary or just below the leaves. Staminate inflorescences 2-8 clustered together, 9-30 by 1-2 mm, puberulous; peduncle absent; bracts broadly triangular, 0.3-0.5 mm long, inconspicuous, glabrous, margin ciliate; glomerules ellipsoid, 1.5-2.2 by c. 1 mm, consisting of more than 10 densely set flowers, approximately continuously set along the rachis or spaced at 0.5-2 mm at base of rachis, continuous apically; pedicel minute. Staminate flowers 0.4-0.8 mm long; sepals 3 or 4, obovate, 0.5-0.7 mm long, almost free, very sparsely puberulous outside, glabrous inside, margin sparsely ciliate; stamens 2, not exserted, 0.3-0.5 mm long; anthers 0.2-0.3 mm long, connective glabrous; pistillode lumpy, minute, ciliate. Pistillate inflorescences 1-3 clustered together, 3-5 by 1-1.2 mm, puberulous, flowers up to 9, densely arranged along the rachis; bracts broadly triangular, 0.3-0.6 mm long, glabrous, margin sparsely ciliate; bracteoles ovate, 0.2-0.4 mm long, sparsely puberulous outside, glabrous inside; pedicel minute, tomentose. Pistillate flowers 2-3.5 mm long; sepals 4 or 5, ovate, patent, 0.6-1 mm long, base fleshy, sparsely hairy at base outside, glabrous inside; ovary ovoid, 1.5-2.5 mm long, 2- (or 3-)locular, punctulate, glabrous or sparsely hairy; stigmas slightly raised, elongated, slightly ascending from the sides of the ovary, straight, apically almost completely divided, 0.3-0.9 mm long, ribbed above, margins papillate to laciniate, glabrous, smooth beneath, glabrous, style remnant present. Infructescences 3-9 by 0.8-1.5 mm, puberulous; fruiting pedicel 0-0.5 mm long, puberulous. Fruits globose to broadly ovoid or ellipsoid, not stiped, sometimes slightly beaked, 7-11 by 6-12 mm, punctulate to finely granular, drying (dark) brown to black, glabrous; pericarp 0.3-0.8 mm thick, not fleshy, taste recorded sweet; septae and column sparsely hirsute at (central) margins (sometimes glabrescent!). Seeds 1, 2 (or 3), half-terete, ellipsoid, 5-8 by 6-7 by 3-4 mm; aril juicy, sweetish. [from Flora Malesiana: http://www.nationaalherbarium.nl/Euphorbs/specA/AporosaT.htm]

Ecology
Primary, secondary, (somewhat) disturbed, and logged over forest, evergreen, mixed dipterocarp, heath or transition between heath and kerangas, submontane forest, fresh water swamp, or periodically inundated; on steep to gentle ridges, hill sides, and hill tops; on flat or (low) undulating land, open, partly shaded, or dry country. Soil: shale, sandy, granitic sand, clayey sand, or clay, yellow or red. Altitude: 0-1200 m.

Distribution
Peninsular Malaysia, Sumatra, Java, Borneo, Sulawesi, Moluccas.

Local names
Borneo Janggau, Kapit-kapit, Kayu massam, Kunau-kunau, Ombah sampori, Penatan, Pupoh, Ubah bunkan.
Sumatra Bangka: Tulang kera.