Aporosa granularis Airy Shaw, Kew Bull. 29 (1974)

Latin for 'knobbled'.

Synonyms
Aporosa microstachya (Tul.) Mull.Arg.

Diagnostics
Shrub up to 9 m tall and 12 cm dbh. Stipules ca. 7 mm long, curved. Leaves alternate, simple, penni-veined, glabrous. Flowers ca. 0.3 mm in diameter, yellowish, placed in racemes. Fruits ca. 8 mm long, yellow-orange-red, dehiscent capsule, seeds with orange aril.

Description
Shrub or tree, up to 9 m high, up to 12 cm diameter. Bark greyish or light brown, smooth to rugose; inner bark pale brown. Young branches (pale) brown, (sparsely) tomentose. Stipules narrowly ovate, slightly oblique, 6-8 by 1.5-2.5 mm, very sparsely sericeous beneath, present in young stages. Petiole terete, adaxially lowly grooved, 7-12 by 0.5-1 mm, tomentose, glabrescent, lower pulvinus 1-1.5 by c. 1 mm, indistinct, upper pulvinus 2-3 by 1-1.2 mm, rather distinct. Leaves narrowly elliptic, 6-16 by 1.8-5 cm; base obtuse to attenuate, basal glands absent; margin very lowly undulate to lowly glandular-crenate, marginal glands few, small, grey, indistinct; apex cuspidate to caudate; blade thin, smooth, dull to slightly shiny, not brittle, drying (dark) greenish grey to greenish brown above, greyish brown beneath, only midrib and nerves puberulous above, glabrescent, sparsely puberulous beneath, midrib and nerves puberulous, glabrescent; dots irregularly set, fine, blackish, mostly indistinct; disc-like glands absent. Nervation: midrib slightly prominent above, prominent beneath; nerves 6-8 pairs, flat to slightly sunken above, prominent beneath, marginal arches distinct, 1-3 mm from the margin; tertiary veins and venation distinct, laxly reticulate, slightly scalariform, flat above, (slightly) prominent beneath. Inflorescences axillary or just below the leaves. Staminate inflorescences 1-3 clustered together, 5-9 by 1-1.5 mm, puberulous; peduncle 0.5-1.5 by c. 0.8 mm; bracts broadly triangular, inconspicuous, 0.3-0.5 mm long, very sparsely puberulous outside, glabrous inside, margin ciliate; glomerules subglobose, c. 1 by 0.5 mm, indistinct, consisting of 3-5 laxly set flowers, some spaced at 0.1-0.5 mm at base of rachis, mostly continuous at apex; pedicel minute. Staminate flowers 0.3-0.5 mm long; sepals 4, obovate, connate at base, 0.4-0.6 mm long, sparsely hairy at base only outside, glabrous inside, margin ciliate; stamens 2, slightly exserted, 0.5-0.7 mm long; anthers 0.1-0.2 mm long; pistillode minute, lumpy. Pistillate inflorescences 1 or 2 clustered together, 2-4 by 0.5-0.8 mm, (sparsely) puberulous; flowers up to 7, mostly densely set at apical part of the rachis; bracts broadly triangular, 0.3-0.5 mm long, sparsely hairy outside, glabrous inside; bracteoles ovate, c. 0.3 mm long, sparsely hairy outside, glabrous inside; pedicel absent. Pistillate flowers 1.5-2 mm long; sepals 4, ovate, patent, 0.7-1 mm long, thin, sparsely hairy outside, glabrous inside; ovary ellipsoid, 2-locular, 1.5-2 mm long, smooth, sparsely sericeous at base, glabrescent; stigmas sessile, elongated, flattened to the sides of the ovary, apically bifid to near base, straight, 1-2 mm long, lowly papillate to laciniate above, glabrous, smooth beneath, glabrous, style remnant absent. Infructescences 3-5 by 0.5-1 mm, sparsely puberulous; fruiting pedicel absent. Fruits ellipsoid, not stiped, not beaked, 7-9 by 6-8 mm, granular, drying (dark) brownish, very sparsely hairy, glabrescent; pericarp 0.3-0.8 mm thick, not fleshy; septae and column glabrous. Seeds 1, half-terete, ellipsoid, 6-7 by 4-5 by 2-3 mm. [from Flora Malesiana: http://www.nationaalherbarium.nl/Euphorbs/specA/AporosaT.htm]

Ecology
Primary, disturbed, old secondary mixed dipterocarp lowland forest, peat swamp, kerangas; on undulating country, steep or gentle slopes, and along river banks. Soil: sandstone or sandy (clay), yellow, rich. Altitude: 50-700 m.

Distribution
Borneo.

Local names
Borneo Ganjii, Janggau, Kayu massam, Kayu matalio.