Ficus glandulifera (Wall. ex Miq.) King, Sp. Ficus 2 (1888)

Latin for 'covered with glands', referring to the glands on the petiole.

Synonyms
Ficus aurantiaca (Miq.) Miq.
Ficus glandulifera var. villosa Corner
Ficus hasskarlii Merr.
Ficus henschelii Merr.
Pogonotrophe aurantiaca Miq.
Pogonotrophe glandulifera Wall. ex Miq.
Pogonotrophe sumatrana Miq.

Diagnostics
Mid-canopy tree up to 33 m tall and 44 cm dbh. Stem with white sap. Stipules ca. 7 mm long, hairy. Petiole with subnodal gland. Leaves alternate, simple, tripli-veined, hairy below. Fruits ca. 10 mm diameter, yellow-orange-red, fleshy figs, placed along the twigs.

Description
Dioecious tree, up to 33 m tall, 44 cm diameter; buttresses up to 1.5 m high. Bark dark brown, smooth and lenticellate; inner bark pale yellow. Sapwood yellowish. Twigs 0.2-0.4 cm thick, reddish brown. Stipules ovate, c. 0.5 cm long, densely reddish brown hairy. Leaves spirally arranged, membranous, densely or sparsely brownish hairy below, very sparsely hairy on midrib and veins above; elliptic to narrowly obovate, (5-)8-15.5 x (2.5-)3.5-8 cm, base cuneate, margin entire, plane, apex cuspidate or acuminate; midrib flat above; lateral veins 4-6 pairs, raised below, faint above, basal pair reaching almost 1/2 the length of the blade, without glands in the axils; intercostal venation scalariform, distinct below, faint above; petiole 1.5-7 cm long, sparsely reddish brown hairy, usually with a subnodal gland. Syconia borne on twigs behind the leaves, ripening yellow to red, subglobose or ellipsoid, c. 1 cm diameter, apex slightly convex and perforated, base abruptly narrowed into a short stalk-like structure; peduncle up to 0.6 cm long; basal bracts 3, 1.5-2.5 mm long; internal bristles numerous, 1-2 mm long, yellowish. Tepals 3-5, narrowly spathulate. Achenes 1-1.5 mm long, rugose, strongly keeled. [from Tree Flora of Sabah and Sarawak]

Ecology
In Undisturbed to slightly disturbed or open mixed dipterocarp, swamp, coastal and sub-montane forests up to 1600 m altitude. Common on alluvial sites, hillsides and ridges. On sandy to clay soils.

Distribution
Thailand, Peninsular Malaysia, Sumatra, Java, Borneo, Philippines, Celebes, Moluccas, New Guinea.

Local names
Borneo: Karai, Lemak-lemak, Lenkan mang, Tanggilam, Tenpan.