Mallotus muticus (Mull.Arg.) Airy Shaw, Kew Bull. 16 (1963)

Latin for 'mute or truncated'.

Coccoceras borneense J.J.Sm.
Coccoceras muticum Mull.Arg.
Coccoceras muticum var. muticum Mull.Arg.
Mallotus borneensis Mull.Arg.
Rottlera borneensis (Mull.Arg.) Scheff.

Upper canopy tree up to 42 m tall and 100 cm dbh. Stipules dropped early. Leaves alternate, simple, tripli-veined, glabrous, usually with several basal nectary glands on upper surface. Flowers ca. 4 mm diameter, green-yellowish, placed in racemes. Fruits ca. 20 mm diameter, reddish, 3-lobed, strongly keeled, indehiscent capsules.

Tree up to 35(-42) m high, dioecious, mainly glabrous; branchlets finely ribbed, not to rarely lenticellate, glabrous, with white to greyish bark with conspicuous discoid glands. Stipules early caducous. Leaves subopposite to alternate, pairs unequal in size; petiole terete to angular, adaxially channelled, 18-83 by 0.7-3 mm, glabrous to sometimes puberulous at ends, base and apex usually not pulvinate; blade elliptic to obovate, 8.2-24.8 by 3.7-14 cm, index 1.3-2.4, chartaceous to coriaceous, base rounded to obtuse and often minutely auriculate, margin entire to slightly irregularly crenate, 6-16 marginal glands per leaf side, usually in sinuses, triangular to globose, thickened, c. 0.7 by 0.3 mm, protruding especially in young leaves, apex acuminate to obtusely cuspidate, sometimes ending in a margin-type of gland, upper surface green to grey to brown, 0-3(-8) basal glands on each side, elliptic, impressed, c. 1.5 by 0.8 mm, grey or brown, 0-8 mm from petiole attachment, apically none to 5 circular, impressed glands on last pair of nerves, lower surface pale green to brown, domatia with villose to long stellate white hairs in nerve axils; venation basally triplinerved, basal nerves ending in margin below and above middle, further pinnate, with 3-5(-7) nerves per side, eucamptodromous, veins conspicuously scalariform. Staminate inflorescences and flowers as those of Mallotus leucodermis, see there. Pistillate inflorescences axillary to rarely terminal, solitary or 2 or 3 together, rachis angular, 3.5-23.5 cm by 0.5-2 mm, 10-35 nodes per rachis, 1 flower per node; bracts 1-1.5 by c. 1 mm, margin glabrous to sometimes ciliate, apex acuminate to cuspidate, patent to reflexed; pedicels terete and often angular near abscission zone, 1-7(-12) by 0.5-1 mm, often geniculate, abscission zone 1-2 mm from apex; flowers 3-4 mm in diam., calyx lobes (4 or) 5, connate to up 0.5 mm from base, triangular to narrowly triangular, 2-2.5 by 0.5-1.2 mm, long white villose hairs on the inside, reddish brown, margin sometimes villosely ciliate, recurved when old, apically acuminate to cuspidate, basally thickened, ovary ovate in lateral view, 2- or 3-locular, covered with discoid glands, style 1.5-2 by c. 0.5 mm, stigmas 2-2.5 by 0.5-1 mm, basally erect, apically recurving. Fruits indehiscent, woody capsules, broadly angular-ovate to obcordate in lateral view, 12-25 by 11-22 mm in diam., apically style remnant often bent; locules 0.7-1 cm wide, slightly to clearly ridged over valve joints; column narrowly rhomboid with basal part slightly longer than apical one, c. 15 mm long by 6 mm wide. Seeds ovoid, 6-7 by 4-5 mm in diam., shiny to dull, smooth, brown. [from Flora Malesiana]

Primary and secondary forest, in periodically inundated forest, forest edges, logged plains, often in riparian forests, and swampy areas. Alt.: sea level to 500 m.

Peninsular Malaysia, Sumatra, Borneo.

Local names
Borneo: Belanti, Kelepuek, Perupuk, Randang jangun, Salung api, Salungapit, Sandanaap, Suko lapit, Tapai logan.
Kalimantan: Kelepuek, perupuk (Modang Dayak).
Sabah: Randang jangun, salungapit or suko lapit (Kedayan); salung api, sandanaap; tapai longan (Murut).
Sumatra: Senggawan.