Mallotus muticus (Mull.Arg.) Airy Shaw, Kew Bull. 16 (1963)
Latin for 'mute or truncated'.
Coccoceras borneense J.J.Sm.
Coccoceras muticum Mull.Arg.
Coccoceras muticum var. muticum Mull.Arg.
Mallotus borneensis Mull.Arg.
Rottlera borneensis (Mull.Arg.) Scheff.
Upper canopy tree up to 42 m tall and 100 cm dbh. Stipules dropped early.
Leaves alternate, simple, tripli-veined, glabrous, usually with several basal
nectary glands on upper surface. Flowers ca. 4 mm diameter, green-yellowish,
placed in racemes. Fruits ca. 20 mm diameter, reddish, 3-lobed, strongly keeled,
Tree up to 35(-42) m high, dioecious, mainly glabrous; branchlets finely ribbed, not to rarely
lenticellate, glabrous, with white to greyish bark with conspicuous discoid glands. Stipules
early caducous. Leaves subopposite to alternate, pairs unequal in size; petiole terete to
angular, adaxially channelled, 18-83 by 0.7-3 mm, glabrous to sometimes puberulous at ends,
base and apex usually not pulvinate; blade elliptic to obovate, 8.2-24.8 by 3.7-14 cm, index
1.3-2.4, chartaceous to coriaceous, base rounded to obtuse and often minutely auriculate,
margin entire to slightly irregularly crenate, 6-16 marginal glands per leaf side, usually
in sinuses, triangular to globose, thickened, c. 0.7 by 0.3 mm, protruding especially in
young leaves, apex acuminate to obtusely cuspidate, sometimes ending in a margin-type of gland,
upper surface green to grey to brown, 0-3(-8) basal glands on each side, elliptic, impressed,
c. 1.5 by 0.8 mm, grey or brown, 0-8 mm from petiole attachment, apically none to 5 circular,
impressed glands on last pair of nerves, lower surface pale green to brown, domatia with villose
to long stellate white hairs in nerve axils; venation basally triplinerved, basal nerves ending
in margin below and above middle, further pinnate, with 3-5(-7) nerves per side, eucamptodromous,
veins conspicuously scalariform. Staminate inflorescences and flowers as those of Mallotus
leucodermis, see there. Pistillate inflorescences axillary to rarely terminal, solitary or 2 or
3 together, rachis angular, 3.5-23.5 cm by 0.5-2 mm, 10-35 nodes per rachis, 1 flower per node;
bracts 1-1.5 by c. 1 mm, margin glabrous to sometimes ciliate, apex acuminate to cuspidate, patent
to reflexed; pedicels terete and often angular near abscission zone, 1-7(-12) by 0.5-1 mm, often
geniculate, abscission zone 1-2 mm from apex; flowers 3-4 mm in diam., calyx lobes (4 or) 5,
connate to up 0.5 mm from base, triangular to narrowly triangular, 2-2.5 by 0.5-1.2 mm, long white
villose hairs on the inside, reddish brown, margin sometimes villosely ciliate, recurved when old,
apically acuminate to cuspidate, basally thickened, ovary ovate in lateral view, 2- or 3-locular,
covered with discoid glands, style 1.5-2 by c. 0.5 mm, stigmas 2-2.5 by 0.5-1 mm, basally erect,
apically recurving. Fruits indehiscent, woody capsules, broadly angular-ovate to obcordate in
lateral view, 12-25 by 11-22 mm in diam., apically style remnant often bent; locules 0.7-1 cm wide,
slightly to clearly ridged over valve joints; column narrowly rhomboid with basal part slightly
longer than apical one, c. 15 mm long by 6 mm wide. Seeds ovoid, 6-7 by 4-5 mm in diam., shiny to
dull, smooth, brown. [from Flora Malesiana]
Primary and secondary forest, in periodically inundated forest, forest edges, logged plains,
often in riparian forests, and swampy areas. Alt.: sea level to 500 m.
Peninsular Malaysia, Sumatra, Borneo.
Borneo: Belanti, Kelepuek, Perupuk, Randang jangun, Salung api, Salungapit,
Sandanaap, Suko lapit, Tapai logan.
Kalimantan: Kelepuek, perupuk (Modang Dayak).
Sabah: Randang jangun, salungapit or suko lapit (Kedayan); salung api, sandanaap; tapai longan