Shorea parvistipulata Heim, Bull. Mens. Soc. Linn. Paris 2 (1891)

Synonyms
Shorea cristata Brandis
Shorea nebulosa Meijer

Diagnostics
Emergent tree up to 70 m tall and 112 cm dbh. Stem with resin. Stipules up to ca. 14 mm long. Leaves alternate, simple, penni-veined, petiole and lower surface with scales, feeling like sand paper, lower surface often slightly glaucous. Flowers ca. 14 mm in diameter, yellow-pink, placed in panicles. Fruits ca. 21 mm long, green-pink, with three ca. 106 mm long wings, wind dispersed.

Description
Very large thinly scaly-barked buttressed tree. Leaf bud, stipule outside, twig, petiole and leaf nervation beneath shortly persistently pale yellow to grey-brown scabrid tomentose, stipule inside and midrib above evenly, not scabridly, so. Twig c. 2 mm diameter, terete, slender, becoming smooth; stipule scars short, pale, prominent, horizontal or descending. Bud to 6 by 4 mm, ovoid, acute. Stipule to 14 by 7 mm, ovate, subacute. Leaves 6-20 by 3-9 cm, very variable in size and shape, +/- oblong, chartaceous; base typically cordate, sometimes broadly cuneate or obtuse: acumen to 1 cm long; nerves 13-21 pairs, usually slender, at 50-80 degrees except at the base, without secondary nerves: tertiary nerves slender, densely scalariform; midrib evident but applanate to depressed above, prominent beneath: petiole 12-15 mm long, short, terete. Panicle to 16 cm long, terminal or axillary, terete, lax, straight, ascending, densely shortly persistently pale brown scabrid pubescent: singly branched, branchlets to 4 cm long, bearing to 11 distichous flowers: bracteoles to 5 by 3 mm, deltoid, acute, puberulent, not at first caducous. Flower bud to 7 by 3 mm, lanceolate. Sepals pubescent on parts exposed in bud: 3 outer narrowly deltoid, acute, 2 inner ovate, narrowly acuminate, shorter and thinner at margins than outer 3. Petals cream sutfused with pink, at base, lanceolate, pubescent on parts exposed in bud. Stamens 15, in 3 unequal verticils: filaments broad and compressed at base, tapering abruptly and filiform beneath the anthers: appendage to connective slender, glabrous, about twice length of anthers. Ovary ovoid, densely pubescent: style filiform, about as long as ovary, glabrous but for a pubescent ring at base. Fruit pedicel to 2 mm long, to 3 mm diameter. Calyx puberulent towards base, glabrescent distally: 3 longer lobes to 20 by 2.5 cm, lorate-spatulate, obtuse. c. 11 mm broad above the to 14 by 18 mm broadly ovate saccate thickened base: 2 shorter lobes to 8 by 0.4 cm, linear, similar at base. Nut to 2.5 by 2 cm, ovoid, frequently crowned by a persistent filiform style remnant, densely evenly buff pubescent. [from Flora Malesiana]

Ecology
In undisturbed mixed dipterocarp forests up to 1500 m altitude. Mostly on hillsides and ridges with clay soils, rarely on sand soils.

Uses
The timber is used.

Distribution
Borneo.

Local names
Borneo: Engkabanh pinang, Engkabang tukul, Kenuar kapas, Meranti, Rangu, Seraya lupa, Seraya merah, Tahan lutup, Tengerawan kuning.