Dacryodes costata (Benn.) H.J. Lam, Ann. Jard. Bot. Btzg. 42 (1932)

Synonyms
Canarium costatum Ridl.
Santiria costata Benn. in Hook.f.

Diagnostics
Mid-canopy tree up to 40 m tall and 72 cm dbh. Leaves alternate, compound, penni-veined, conspicuously thickened petiole base and tips, petiole hairy. Flowers ca. 5 mm in diameter, yellowish, placed in panicles. Fruits ca. 15 mm long, yellow-orange, fleshy drupe.

Description
Tree 20-30(-40) m by 20-100 cm, rarely with small buttresses. Branchlets c. 0.5 cm thick, minutely pubescent; pith without vascular strands. Leaves (1-)2(-4)-jugate, more or less densely pubescent except the upper surface of the leaflets. Petioles 1-5(-6.5) cm, much flattened at base, pith with few vascular strands. Leaflets broadly elliptic to oblong, 3-16 by 1.5-7.5 cm, rigid, chartaceous to subcoriaceous, underneath more or less woolly pubescent on midrib and nerves, glabrescent; base hardly oblique, broadly cuneate to rounded; apex abruptly, shortly, and bluntly acuminate; nervation prominent beneath, nerves 6-13 pairs (angle c. 60 degrees, at the base up to 90 degrees), strongly curved, not arching except some apical ones. Panicles terminal and often in the uppermost leaf-axils, usually branched from the base, densely woolly pubescent, flowers in clusters; male ones 6.5-30 cm, much branched, branches spreading, up to 12 cm; female ones 4-21 cm, narrowly paniculate to subracemose, branches up to 4 cm. Flowers small, slightly pubescent. Calyx 0.5-1 mm high. Petals glabrous. Filaments free from the disk. Disk annular to 6-lobed, thick. Pistil in male flowers very reduced. Infructescences densely woolly pubescent. Fruits ovoid, hardly or not oblique, 12-19 by 8-12 mm, rounded at the base, more or less obtusely acute at the apex; wall of the pyrene rather thick and hard. Cotyledons folded, 3-lobed. [from Flora Malesiana]

Ecology
In undisturbed mixed dipterocarp forests up to 700 m altitude. Usually on hillsides and ridges with sandy soils.

Uses
The timber is used for light construction, the bole for building canoes.

Distribution
Peninsular Malaysia, Sumatra, Borneo, Philippines.

Local names
Bangka: binjau, bunjau, sekibai, sudur bajan.
Biliton: berbing pinggan.
Borneo: Batu, Basi-basi, Belanak, Buah, Kedongdong, Kramu, Kumbajau, Limat, Unggit.
Malaysia: kedondong (mata-hari).
Sumatra: Kedondong besi, kening kerak, merasam pujo, rasak babi, resting, sepah (or sepat) burung, sikatjang, tjak sibui, tjetjibui.