Glochidion littorale Blume, Bijdr. (1825)
Latin for 'coastal'.
Agyneia impuber Wall. [Invalid]
Bradleia littorea (Blume) Steud.
Bradleia obtusa Wall.
Diasperus litoralis (Blume) Kuntze
Glochidion littorale var. caudatum Airy Shaw
Glochidion littorale var. culminicola Airy Shaw
Phyllanthus littoralis (Blume) M¨šll.Arg.
Shrub up to 10 m tall and 15 cm dbh. Stipules ca. 1.5 mm long, lost early. Leaves
alternate, simple, penni-veined, glabrous, roundish, thick leathery. Flowers ca. 4 mm diameter,
green-white-yellowish, placed in axillary bundles. Fruits ca. 12 mm long,
pink-reddish, dehiscent capsules, seeds with orange-red arils.
Shrubs up to 10 m high; flowering twigs 1.5-2 mm thick; glabrous. Leaves: stipules triangular,
1.3-1.8 by 0.7-2 mm, caducous; petiole 3-4 mm long; blade elliptic to obovate, 2.2-9.2 by 1.9-5 cm,
length/width ratio 1.2-1.8, coriaceous, base (rounded to) shortly attenuate, slightly asymmetric,
margin flat, apex emarginate to rounded, often drying greenish above, brown underneath; venation
slightly raised above, raised below, nerves 7 or 8 per side, sometimes yellow, veins coarsely
reticulate. Inflorescences fascicles of up to 5 flowers, two or three of which pistillate. Flowers:
sepals 6, basally united, margins usually hyaline. Staminate flowers 4.5-5.5 mm in diameter; pedicel
5.5-6 mm long, light green; calyx light green, tube 0.8-1 mm long, lobes elliptic to obovate, outer
3.6-4.2 by 1.8-1.9 mm, inner 3.2-3.4 by 1.5-1.6, usually folded, apically somewhat hooded; stamens
5 or 6, androecium c. 1.4 mm long, stamens c. 1.1 mm long, connective teeth 0.2-0.3 mm long.
Pistillate flowers 3-3.5 mm in diameter; pedicel 2-6.5 mm in fruit; calyx outer lobes triangular
to elliptic, 1.3-2.3 by 1.4-2 mm, inner lobes elliptic 1.3-2.7 by 0.6-2.7 mm; ovary 10-14-locular,
c. 2.7 by 2.3 mm high, longitudinally ribbed; ovules superposed; stigmas in a cone, c. 2 mm long,
11-14-dentate. Fruits round, apically flattened and depressed in centre, basally sometimes also
flattened, 1.4-1.8 by 0.8-1.3 cm high, not lobed, sutures depressed, pinkish-whitish to dark pink;
wall woody, up to 1 mm thick; column c. 12 mm long, spindle-shaped. Seeds c. 6 by 4.5 mm (a single
ovule developing per locule) to 2.5-3.5 by c. 4.5 mm (both ovules developing), red. [from Flora
Coastal or tidal regions with open (disturbed) vegetation types, also in
keranga forest and swamps up to 100 m altitude. Usually on poor sandy soils.
The root is used medicinal as mouthwash for babies. The fruits are edible.
India, Sri Lanka, Thailand, Peninsular Malaysia, Sumatra, Java, Borneo, Philippines, Celebes.
Borneo: Kabassi-kabassi, Obar-obar, Sakah-sakah, Sakah-sakah bini.
Thailand: Nok non thale (Narathiwat), Man Pu (Peninsular).