Mallotus spinifructus Welzen & SEC Sierra, Blumea 51: 382 (2006)(
Name meaning 'spiny fruits'.
Small trees. Stipules present but dropped early. Leaves opposite, with one of each pair smaller.
Petiole swollen basally and apically. Leaf blade with two nectary glands at the base, margin
Trees, up to 10 m high, dbh up to 12 cm; flowering branchlets 3-4 mm thick, flattened and somewhat
ridged when dry, apical parts stellately hairy, glabrescent; bracts of the terminal buds triangular,
c. 1.5 by 0.7 mm. Outer bark grey to greyish pale brown, firm, smooth; inner bark dull brown,
coarsely fibrous. Indumentum of simple and stiff, stellately bundled hairs, glandular scales
yellowish. Stipules triangular, basally usually quickly decreasing in width, 7-10 by 1-1.7 mm,
rather late caducous, outside hairy. Leaves opposite, similar in shape, one somewhat smaller;
petiole 1-6 cm long, round, hairy, basally and apically pulvinate, especially upper pulvinus
well-developed, oblique with blade base partly overlaying; blade ovate to elliptic, 14.5-24.3 by
5-9.5 cm, length/width ratio 2.6-2.9, symmetric, membranous, base cuneate, very base narrowly
cordate, margin entire, apex caudate, upper surface basally with one pair of somewhat elliptic
extrafloral nectaries, up to 1.1 by 0.9 mm, in upper third usually several small extrafloral
nectaries (mainly 0.3 by 0.3 mm) on the nerves, at most slightly hairy on the midrib; lower surface
hairy on venation, glandular scales distinct, hair tuft domatia present; venation triplinerved,
flat and somewhat indistinct above, raised and distinct beneath, yellowish, 8 or 9 nerves per side,
veins and veinlets scalariform. Inflorescences racemes, axillary to (pseudo)terminal, single,
densely hairy; staminate ones only seen in bud, up to 6 cm long, c. 0.8 mm wide, with groups of up
to c. 10 buds, pistillate inflorescences up to 7.5 cm long, a single flower per node; bract
triangular, 5.5-6 by 1.3-1.8 mm, densely hairy outside, almost floccose, recurved. Staminate
flowers only seen in bud, buds globose, pedicelled; pedicel densely hairy; calyx apparently
splitting into three sepals, densely hairy outside; stamens perhaps c. 30, anthers separated by
connective. Pistillate flowers c. 7.5 mm diam., subsessile when young, in fruit pedicel up to 8? mm
long, hairy; sepals 5, triangular to ovate, 4.5-5 by 1.3-2 mm, hairy outside, reflexed; ovary 3-
(or 4-)locular, densely long spiny, spines not in groups but pointing all directions, with loose
long patent white hairs, glandular scales present; style c. 1.5 mm long, stigmas c. 7 mm long,
c. 1 mm wide, with long, branching papillae above, (sub)glabrous beneath. Fruits lobed capsules,
1.8-1.9 by 1-1.1 cm, green when immature, spines up to 4.5 mm long, very slender; wall woody,
basally c. 0.7 mm thick, apically c. 1.7 mm thick; column broadly T-shaped to obtriangular, c.
5.5 mm long. Seeds subglobose 6-8 by 5-7 mm, shiny.
(Logged) riverine forest, alluvial soil, well-drained. Altitude 200-300 m.
Peninsular Malaysia and Borneo.
Borneo: Kreh merah.